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Homosexual behavior in animals is sexual behavior among non-human species that is interpreted as homosexual or bisexual. This may include same-sex sexual activity , courtship , affection , pair bonding , and parenting among same-sex animal pairs. The sexual behavior of non-human animals takes many different forms, even within the same species, though homosexual behavior is best known from social species.

Scientists perceive homosexual behavior in animals to different degrees. The motivations for and implications of these behaviors have yet to be fully understood, since most species have yet to be fully studied. Thus, a homosexual orientation, if one can speak of such thing in animals, seems to be a rarity.

According to Bagemihl , same-sex behavior comprising courtship, sexual, pair-bonding, and parental activities has been documented in over species of animals worldwide. The term homosexual was coined by Karl-Maria Kertbeny in to describe same-sex sexual attraction and sexual behavior in humans. According to Bruce Bagemihl , when describing animals, the term homosexual is preferred over gay , lesbian , and other terms currently in use, as these are seen as even more bound to human homosexuality.

In humans, the term is used to describe individual sexual behaviors as well as long-term relationships, but in some usages connotes a gay or lesbian social identity. Scientific writing would benefit from reserving this anthropomorphic term for humans and not using it to describe behavior in other animals, because of its deeply rooted context in human society".

Animal preference and motivation is always inferred from behavior. In wild animals, researchers will as a rule not be able to map the entire life of an individual, and must infer from frequency of single observations of behavior. In most instances, it is presumed that the homosexual behavior is but part of the animal's overall sexual behavioral repertoire, making the animal "bisexual" rather than "homosexual" as the terms are commonly understood in humans.

The observation of homosexual behavior in animals can be seen as both an argument for and against the acceptance of homosexuality in humans, and has been used especially against the claim that it is a peccatum contra naturam "sin against nature". For instance, homosexuality in animals was cited by the American Psychiatric Association and other groups in their amici curiae brief to the United States Supreme Court in Lawrence v.

Texas , which ultimately struck down the sodomy laws of 14 states. A majority of the research available concerning homosexual behavior in animals lacks specification between animals that exclusively exhibit same-sex tendencies and those that participate in heterosexual and homosexual mating activities interchangeably.

This lack of distinction has led to differing opinions and conflicting interpretations of collected data amongst scientists and researchers. For instance, Bruce Bagemihl , author of the book Biological Exuberence: Animal Homosexuality and Natural Diversity , emphasizes that there are no anatomical or endocrinological differences between exclusively homosexual and exclusively heterosexual animal pairs.

Many of the animals used in laboratory-based studies of homosexuality do not appear to spontaneously exhibit these tendencies often in the wild.

Such behavior is often elicited and exaggerated by the researcher during experimentation through the destruction of a portion of brain tissue, or by exposing the animal to high levels of steroid hormones prenatally.

Homosexual behaviour in animals has been discussed since classical antiquity. The earliest written mention of animal homosexuality appears to date back to 2, years ago, when Aristotle — BC described copulation between pigeons, partridges and quails of the same sex. Until recent times, the presence of same-sex sexual behavior was not "officially" observed on a large scale, possibly due to observer bias caused by social attitudes to same-sex sexual behavior, [27] innocent confusion, lack of interest, distaste, scientists fearing loss of their grants or even from a fear of "being ridiculed by their colleagues".

The true extent of homosexuality in animals is not known. No species has been found in which homosexual behaviour has not been shown to exist, with the exception of species that never have sex at all, such as sea urchins and aphis. Moreover, a part of the animal kingdom is hermaphroditic, truly bisexual. For them, homosexuality is not an issue. An example of overlooking homosexual behavior is noted by Bagemihl describing mating giraffes where nine out of ten pairings occur between males:.

Every male that sniffed a female was reported as sex, while anal intercourse with orgasm between males was only "revolving around" dominance, competition or greetings.

Some researchers believe this behavior to have its origin in male social organization and social dominance, similar to the dominance traits shown in prison sexuality. Others have argued that social organization theory is inadequate because it cannot account for some homosexual behaviors, for example, penguin species where male individuals mate for life and refuse to pair with females when given the chance.

In fact, apparent homosexual individuals are known from all of the traditional domestic species, from sheep, cattle and horses to cats, dogs and budgerigars. A definite physiological explanation or reason for homosexual activity in animal species has not been agreed upon by researchers in the field.

Numerous scholars are of the opinion that varying levels either higher or lower of the sex hormones in the animal, [37] in addition to the size of the animal's gonads, [24] play a direct role in the sexual behavior and preference exhibited by that animal.

Others firmly argue no evidence to support these claims exists when comparing animals of a specific species exhibiting homosexual behavior exclusively and those that do not.

Ultimately, empirical support from comprehensive endocrinological studies exist for both interpretations. Additional studies pertaining to hormone involvement in homosexual behavior indicate that when administering treatments of testosterone and estradiol to female heterosexual animals, the elevated hormone levels increase the likelihood of homosexual behavior.

Additionally, boosting the levels of sex hormones during an animal's pregnancy appears to increase the likelihood of it birthing a homosexual offspring. Researchers found that disabling the fucose mutarotase FucM gene in laboratory mice — which influences the levels of estrogen to which the brain is exposed — caused the female mice to behave as if they were male as they grew up.

However, in addition to homosexual behavior, several abnormal behaviors were also exhibited apparently due to this mutation. In March , research showed that serotonin is involved in the mechanism of sexual orientation of mice.

An estimated one-quarter of all black swans pairings are of males. They steal nests, or form temporary threesomes with females to obtain eggs, driving away the female after she lays the eggs. The males spent time in each other's society, guarded the common territory, performed greeting ceremonies before each other, and in the reproductive period pre-marital rituals, and if one of the birds tried to sit on the other, an intense fight began.

The same reasoning has been applied to male flamingo pairs raising chicks. Female albatross, on the north-western tip of the island of Oahu, Hawaii, form pairs for co-growing offspring. Research has shown that the environmental pollutant methylmercury can increase the prevalence of homosexual behavior in male American white ibis. The study involved exposing chicks in varying dosages to the chemical and measuring the degree of homosexual behavior in adulthood.

The results discovered was that as the dosage was increased the likelihood of homosexual behavior also increased. The endocrine blocking feature of mercury has been suggested as a possible cause of sexual disruption in other bird species. Mallards form male-female pairs only until the female lays eggs, at which time the male leaves the female. Penguins have been observed to engage in homosexual behaviour since at least as early as The report was considered too shocking for public release at the time, and was suppressed.

The only copies that were made available privately to researchers were translated into Greek, to prevent this knowledge becoming more widely known. The report was unearthed only a century later, and published in Polar Record in June In early February the New York Times reported that Roy and Silo , a male pair of chinstrap penguins in the Central Park Zoo in New York City had successfully hatched and fostered a female chick from a fertile egg they had been given to incubate.

In Odense Zoo in Denmark, a pair of male king penguins adopted an egg that had been abandoned by a female, proceeding to incubate it and raise the chick. Researchers at Rikkyo University in Tokyo found 20 homosexual pairs at 16 major aquariums and zoos in Japan. The Bremerhaven Zoo in Germany attempted to encourage reproduction of endangered Humboldt penguins by importing females from Sweden and separating three male pairs, but this was unsuccessful.

The zoo's director said that the relationships were "too strong" between the homosexual pairs. A pair of male Magellanic penguins who had shared a burrow for six years at the San Francisco Zoo and raised a surrogate chick, split when the male of a pair in the next burrow died and the female sought a new mate.

Buddy and Pedro, a pair of male African penguins , were separated by the Toronto Zoo to mate with female penguins. Chupchikoni was assumed to be male until her blood was tested. In Jumbs and Hurricane, two Humboldt penguins at Wingham Wildlife Park became the center of international media attention as two male penguins who had pair bonded a number of years earlier and then successfully hatched and reared an egg given to them as surrogate parents after the mother abandoned it halfway through incubation.

In two male griffon vultures named Dashik and Yehuda, at the Jerusalem Biblical Zoo , engaged in "open and energetic sex" and built a nest. The keepers provided the couple with an artificial egg, which the two parents took turns incubating; and 45 days later, the zoo replaced the egg with a baby vulture. The two male vultures raised the chick together. Dashik became depressed, and was eventually moved to the zoological research garden at Tel Aviv University where he too set up a nest with a female vulture.

Two male vultures at the Allwetter Zoo in Muenster built a nest together, although they were picked on and their nest materials were often stolen by other vultures.

They were eventually separated to try to promote breeding by placing one of them with female vultures, despite the protests of German homosexual groups. Both male and female pigeons sometimes exhibit homosexual behavior. In addition to sexual behavior, same-sex pigeon pairs will build nests, and hens will lay infertile eggs and attempt to incubate them. The Amazon river dolphin or boto has been reported to form up in bands of 3—5 individuals engaging in sexual activity.

The groups usually comprise young males and sometimes one or two females. Sex is often performed in non-reproductive ways, using snout, flippers and genital rubbing, without regard to gender.

Courtship, mounting, and full anal penetration between bulls has been noted to occur among American bison. The Mandan nation Okipa festival concludes with a ceremonial enactment of this behavior, to "ensure the return of the buffalo in the coming season".

The behaviour is hormone driven and synchronizes with the emergence of estrus heat , particularly in the presence of a bull. More than 20 species of bat have been documented to engage in homosexual behavior.

Bat species that have been observed engaging in homosexual behavior in captivity include the Comoro flying fox Pteropus livingstonii , the Rodrigues flying fox Pteropus rodricensis and the common vampire bat Desmodus rotundus. Homosexual behavior in bats has been categorized into 6 groups: In the wild, the grey-headed flying fox Pteropus poliocephalus engages in allogrooming wherein one partner licks and gently bites the chest and wing membrane of the other partner.

Both sexes display this form of mutual homosexual grooming and it is more common in males. Males often have erect penises while they are mutually grooming each other.

In wild Bonin flying foxes Pteropus pselaphon , males perform fellatio or 'male-male genital licking' on other males. Male—male genital licking events occur repeatedly several times in the same pair, and reciprocal genital licking also occurs. The male-male genital licking in these bats is considered a sexual behavior. Allogrooming in Bonin flying foxes has never been observed, hence the male-male genital licking in this species does not seem to be a by-product of allogrooming, but rather a behavior of directly licking the male genital area, independent of allogrooming.

In wild Indian flying foxes Pteropus giganteus , males often mount one another, with erections and thrusting, while play-wrestling. A similar behavior was also observed in the common bent-wing bat Miniopterus schreibersii. In wild little brown bats Myotis lucifugus , males often mount other males and females during late autumn and winter, when many of the mounted individuals are torpid.

The lethargic males, like females, called out loudly and presented their buccal glands with opened mouth during copulation. Vesey-Fitzgerald observed homosexual behaviours in all 12 British bat species known at the time: I have even seen homosexuality between Natterer's and Daubenton's bats Myotis nattereri and M. Dolphins of several species engage in homosexual acts, though it is best studied in the bottlenose dolphins.

Janet Mann, Georgetown University professor of biology and psychology, argues that the strong personal behavior among male dolphin calves is about bond formation and benefits the species in an evolutionary context.

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